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I don't think I ever planned what its original trait would've been. Maybe make it something related to storms?
In adv12, it makes it clear that Avilona used to be (metaphysically at least) an Eagle. And now the Eagle has died.

So riffing off of that, maybe everyone can see really far? (i.e. the world doesn't need telescopes)

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I'd probably just keep it simple. Original trait: 'long-lasting, high-speed flight is easy.' I seem to recall that there was a thing in the books or that you told me, Ryan, that pre-Skyfall the dragons could fly at hundreds of miles per hour.

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