So this is what the FAQ on DnDBeyond states:
"
What does backward compatible mean?
It means that fifth edition adventures and supplements will work in One D&D. For example, if you want to run Curse of Strahd in One D&D, that book will work with the new versions of the core rulebooks. Our goal is for you to keep enjoying the content you already have and make it even better. You’ll see this in action through the playtest materials, which you will be able to provide feedback on.
"
I see the design commitment is that DMs can use supplements − like Xanathars and Tashas − with the 5.5 core books.
At the same time, these also work with the 5.0 core books.
Presumably, 5.0 and 5.5 can both work at the same table, even if it requires 5.0 getting a free feat or 5.5 using 5.0 backgrounds instead.